Meiosis, which occurs during sexual reproduction, allows for a great variety of possible gene combinations. Which of the following can increase genetic variation during meiosis?

the duplication of a gene caused by an error in chromosomal crossover
the transposition of a gene from one place in an organism's genome to another
the deletion of a gene caused by an error in chromosomal crossover
the exchange of genes during chromosomal crossover

All of the options listed can increase genetic variation during meiosis.