What does the presence of a vestigial structure in a modern organism indicate?

a
The structure was probably functional in some ancestor of the modern organism
b
A limited number of genes are expressed during the lifetime of an organism
c
Natural selection perpares the body of organisms so that in the future the vestigial structures may be used
d
The vestigial structure was not used by the modern organism, so it became nonfunctional within the organism's lifetime

a. The structure was probably functional in some ancestor of the modern organism