what is the function of endodermis in the root

The endodermis is a plant tissue that surrounds the root's vascular tissue (the xylem and phloem). Its function is to regulate the movement of water, ions, and other substances from the soil into the plant's root and up to the rest of the plant. The endodermis achieves this by containing a waxy substance called suberin in its cell walls, which forms an impermeable barrier around each cell. This barrier forces all water and substances to be filtered through the selectively permeable plasma membranes of the endodermal cells to reach the rest of the plant, allowing the endodermis to act as a gatekeeper for the plant's nutrient uptake.

The endodermis is a specialized layer of cells located in the innermost part of the root cortex. Its main function is to regulate the movement of substances into the vascular cylinder, which is the central column of tissues in the root that contains the xylem and phloem.

Here are the step-by-step functions of the endodermis in the root:

1. Protection: The endodermis acts as a protective layer for the inner tissues of the root. It provides a barrier that prevents pathogens, toxins, and excess water from entering the vascular cylinder.

2. Selective Absorption: The endodermis is selectively permeable, meaning it controls the movement of substances into the vascular cylinder. It contains a specialized band called the Casparian strip, which is made up of suberin—a waxy substance that blocks the movement of water and solutes between the cells. This forces nutrients and water to pass through the endodermal cells, enabling their selective absorption and preventing uncontrolled movement.

3. Regulation of Water and Mineral Uptake: The endodermis plays a crucial role in regulating the movement of water and minerals absorbed by the root hairs. By controlling the entry into the vascular cylinder, it helps maintain the proper balance of ions and nutrients within the plant.

4. Control of Hormone Transport: The endodermis also helps regulate the transport of hormones, such as auxins and cytokinins, in the root. These hormones play key roles in the growth and development of the plant, and their movement through the endodermis can be regulated by specialized transport proteins.

5. Root Pressure Maintenance: The endodermis, along with the other cells of the root, helps generate and maintain root pressure. This pressure is essential for the movement of water and minerals up the plant's xylem vessels and contributes to processes like guttation (the exudation of water droplets from the leaf margins).

In summary, the endodermis in the root functions to protect the inner tissues, selectively absorb nutrients and water, regulate hormone transport, and maintain root pressure.