what is the subject of the sentence...Two lost species are the passenger pigeon and the carolina parakeet.

There are three nouns and no pronouns in this sentence. Which noun do you think is the subject?

species
pigeon
parakeet

(If you think that passenger and carolina are nouns, too, you're right in a way, but they are being used as adjectives in this particular sentence.)

Let us know what you think.

http://grammar.ccc.commnet.edu/grammar/subjects.htm

http://www.uottawa.ca/academic/arts/writcent/hypergrammar/rvsubj.html

http://www.ucalgary.ca/UofC/eduweb/grammar/index.htm

=)

To identify the subject of a sentence, you can follow these steps:

1. Identify the verb: In the given sentence, the verb is "are." It helps to note that the subject usually performs the action expressed by the verb.

2. Ask "who" or "what" before the verb: In this case, you can ask "Who or what are?" The answer will give you the subject of the sentence.

Applying these steps to the sentence "Two lost species are the passenger pigeon and the Carolina parakeet," the subject is "Two lost species." The subject is the entity being described as that is what the verb "are" is referring to.